{"id":65923,"date":"2019-02-04T16:12:55","date_gmt":"2019-02-05T00:12:55","guid":{"rendered":"https:\/\/sou.edu\/academics\/?page_id=65923"},"modified":"2025-03-28T16:20:52","modified_gmt":"2025-03-28T23:20:52","slug":"john-roden","status":"publish","type":"page","link":"https:\/\/sou.edu\/academics\/biology\/faculty\/john-roden\/","title":{"rendered":"John S. Roden"},"content":{"rendered":"\n<style type=\"text\/css\" data-created_by=\"avia_inline_auto\" id=\"style-css-av-50sq96w-b1168545cc87b94022f35bf82e8db676\">\n.avia-section.av-50sq96w-b1168545cc87b94022f35bf82e8db676{\nbackground-color:#f3f3f3;\nbackground-image:unset;\n}\n<\/style>\n<div id='av_section_1' aria-label='SOU Biology Individual Faculty Page - John S. Roden Intro Section' class='avia-section av-50sq96w-b1168545cc87b94022f35bf82e8db676 main_color avia-section-default avia-no-border-styling  avia-builder-el-0  el_before_av_submenu  avia-builder-el-first  avia-bg-style-scroll container_wrap fullsize'  ><div class='container av-section-cont-open' ><main  role=\"main\" itemprop=\"mainContentOfPage\"  class='template-page content  av-content-full alpha units'><div class='post-entry post-entry-type-page post-entry-65923'><div class='entry-content-wrapper clearfix'>\n\n<style type=\"text\/css\" data-created_by=\"avia_inline_auto\" id=\"style-css-av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2\">\n#top .av-special-heading.av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2{\npadding-bottom:0;\nfont-size:36px;\n}\nbody .av-special-heading.av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2 .av-special-heading-tag .heading-char{\nfont-size:25px;\n}\n#top #wrap_all .av-special-heading.av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2 .av-special-heading-tag{\nfont-size:36px;\n}\n.av-special-heading.av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2 .av-subheading{\nfont-size:18px;\n}\n\n@media only screen and (min-width: 480px) and (max-width: 767px){ \n#top #wrap_all .av-special-heading.av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2 .av-special-heading-tag{\nfont-size:0.8em;\n}\n}\n\n@media only screen and (max-width: 479px){ \n#top #wrap_all .av-special-heading.av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2 .av-special-heading-tag{\nfont-size:0.8em;\n}\n}\n<\/style>\n<div  class='av-special-heading av-jrtqinu3-0a80ca5bf2fecc2ed9a96c34b1b8b6e2 av-special-heading-h1 blockquote modern-quote modern-centered  avia-builder-el-1  avia-builder-el-no-sibling  av-inherit-size'><h1 class='av-special-heading-tag '  itemprop=\"headline\"  >Biology Faculty<\/h1><div class='av-subheading av-subheading_below'><p>John S. Roden<\/p>\n<\/div><div class=\"special-heading-border\"><div class=\"special-heading-inner-border\"><\/div><\/div><\/div>\n\n<\/div><\/div><\/main><!-- close content main element --><\/div><\/div><div class='clear'><\/div><div id='sub_menu1'  class='av-submenu-container av-jrqyml9g-fa191134e429c2e0328a9b4a1e4a72cf alternate_color  avia-builder-el-2  el_after_av_section  el_before_av_section  no-border undergrad-goofy-foot av-sticky-submenu submenu-not-first container_wrap fullsize' style='z-index:301' ><div class='container av-menu-mobile-disabled av-submenu-pos-center'><ul id=\"menu-biology\" class=\"av-subnav-menu\" role=\"menu\"><li role=\"menuitem\" id=\"menu-item-57387\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-1\"><a href=\"https:\/\/sou.edu\/academics\/biology\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Biology Degree Program<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-57391\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-2\"><a href=\"https:\/\/sou.edu\/academics\/biology\/programs\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Programs<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-87730\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-3\"><a href=\"https:\/\/sou.edu\/academics\/biology\/biology-stories\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Stories<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-67857\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-4\"><a href=\"https:\/\/sou.edu\/academics\/biology\/facilities\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Facilities<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-66579\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-5\"><a href=\"https:\/\/sou.edu\/academics\/biology\/advising\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Advising<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-66577\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-6\"><a href=\"https:\/\/sou.edu\/academics\/biology\/scholarships\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Scholarships<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-57390\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-7\"><a href=\"https:\/\/sou.edu\/academics\/biology\/about\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">About<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-57389\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-8\"><a href=\"https:\/\/sou.edu\/academics\/biology\/faculty\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Faculty<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<li role=\"menuitem\" id=\"menu-item-57388\" class=\"menu-item menu-item-type-post_type menu-item-object-page menu-item-top-level menu-item-top-level-9\"><a href=\"https:\/\/sou.edu\/academics\/biology\/contact\/\" itemprop=\"url\" tabindex=\"0\"><span class=\"avia-bullet\"><\/span><span class=\"avia-menu-text\">Contact<\/span><span class=\"avia-menu-fx\"><span class=\"avia-arrow-wrap\"><span class=\"avia-arrow\"><\/span><\/span><\/span><\/a><\/li>\n<\/ul><\/div><\/div><div class='sticky_placeholder'><\/div>\n<div id='av_section_2' aria-label='SOU Biology Individual Faculty Page - John S. Roden Biography Section' class='avia-section av-1fh2y20-88aad0089b7e1e287224c438418d5aeb main_color avia-section-default avia-shadow  avia-builder-el-3  el_after_av_submenu  el_before_av_section  bio-list avia-bg-style-scroll container_wrap fullsize'  ><div class='container av-section-cont-open' ><div class='template-page content  av-content-full alpha units'><div class='post-entry post-entry-type-page post-entry-65923'><div class='entry-content-wrapper clearfix'>\n\n<style type=\"text\/css\" data-created_by=\"avia_inline_auto\" id=\"style-css-av-11guxu0-b229c518ee7a3c1fe45fa23e1dafe556\">\n.flex_column.av-11guxu0-b229c518ee7a3c1fe45fa23e1dafe556{\nborder-radius:0px 0px 0px 0px;\npadding:0px 0px 0px 0px;\n}\n<\/style>\n<div  class='flex_column av-11guxu0-b229c518ee7a3c1fe45fa23e1dafe556 av_one_fourth  avia-builder-el-4  el_before_av_three_fourth  avia-builder-el-first  first flex_column_div av-zero-column-padding  '     ><style type=\"text\/css\" data-created_by=\"avia_inline_auto\" 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https:\/\/sou.edu\/academics\/wp-content\/uploads\/sites\/14\/2018\/09\/John-Roden-Southern-Oregon-University-Biology-Faculty-36x36.png 36w, https:\/\/sou.edu\/academics\/wp-content\/uploads\/sites\/14\/2018\/09\/John-Roden-Southern-Oregon-University-Biology-Faculty-180x180.png 180w, https:\/\/sou.edu\/academics\/wp-content\/uploads\/sites\/14\/2018\/09\/John-Roden-Southern-Oregon-University-Biology-Faculty-120x120.png 120w\" sizes=\"(max-width: 183px) 100vw, 183px\" class=\"wp-image-64008 avia-img-lazy-loading-not-64008 avia_image  sp-no-webp\" alt=\"John Roden 麻豆传媒女神 Biology Faculty\" decoding=\"async\" fetchpriority=\"high\" itemprop=\"thumbnailUrl\"  > <\/picture><\/div><\/div><\/div><\/div>\n<style type=\"text\/css\" data-created_by=\"avia_inline_auto\" id=\"style-css-av-n65pd4-43d7c8ea14e100ab4383b7e07140d702\">\n.flex_column.av-n65pd4-43d7c8ea14e100ab4383b7e07140d702{\nborder-radius:0px 0px 0px 0px;\npadding:0px 0px 0px 0px;\n}\n<\/style>\n<div  class='flex_column av-n65pd4-43d7c8ea14e100ab4383b7e07140d702 av_three_fourth  avia-builder-el-6  el_after_av_one_fourth  avia-builder-el-last  flex_column_div av-zero-column-padding  '     ><p>\n<style type=\"text\/css\" data-created_by=\"avia_inline_auto\" id=\"style-css-av-176qmub-25b58d9907d220235bca6a91175ac841\">\n#top .av-special-heading.av-176qmub-25b58d9907d220235bca6a91175ac841{\npadding-bottom:0;\nfont-size:20px;\n}\nbody .av-special-heading.av-176qmub-25b58d9907d220235bca6a91175ac841 .av-special-heading-tag .heading-char{\nfont-size:25px;\n}\n#top #wrap_all .av-special-heading.av-176qmub-25b58d9907d220235bca6a91175ac841 .av-special-heading-tag{\nfont-size:20px;\n}\n.av-special-heading.av-176qmub-25b58d9907d220235bca6a91175ac841 .av-subheading{\nfont-size:15px;\n}\n\n@media only screen and (min-width: 480px) and (max-width: 767px){ \n#top #wrap_all .av-special-heading.av-176qmub-25b58d9907d220235bca6a91175ac841 .av-special-heading-tag{\nfont-size:0.8em;\n}\n}\n\n@media only screen and (max-width: 479px){ \n#top #wrap_all .av-special-heading.av-176qmub-25b58d9907d220235bca6a91175ac841 .av-special-heading-tag{\nfont-size:0.8em;\n}\n}\n<\/style>\n<div  class='av-special-heading av-176qmub-25b58d9907d220235bca6a91175ac841 av-special-heading-h4 blockquote modern-quote  avia-builder-el-7  el_before_av_textblock  avia-builder-el-first  av-inherit-size av-linked-heading'><h4 class='av-special-heading-tag '  itemprop=\"headline\"  >John S. Roden<\/h4><div class=\"special-heading-border\"><div class=\"special-heading-inner-border\"><\/div><\/div><\/div><br \/>\n<section  class='av_textblock_section av-jrqymfzv-156feb6f52d13c9d1949d348e849593f '   itemscope=\"itemscope\" itemtype=\"https:\/\/schema.org\/CreativeWork\" ><div class='avia_textblock'  itemprop=\"text\" ><p class=\"pop-4 dig-4 bio-cred\"><em>Associate Professor<\/em><br \/>\nPhD, Botany, University of California, Davis, 1992; MS, Botany, University of California, Davis, 1989; BS, Forest Management, University of Washington, 1987<\/p>\n<p class=\"dig-4\"><span class=\"heavy-man\">Teaching:<\/span><\/p>\n<ul class=\"pop-4\">\n<li>Plant Physiology (Bi 331)<\/li>\n<li>Principles of Biology (Bi211)<\/li>\n<li>Forest Ecology and Management (Bi386)<\/li>\n<li>Biogeochemistry (Bi346)<\/li>\n<li>Plant Form and Function (Bi434\/534)<\/li>\n<li>Introductory Ecology Lab (Bi340L)<\/li>\n<li>Stable Isotope Ecology and Biogeochemistry (summer course, University of Utah)<\/li>\n<\/ul>\n<p class=\"dig-4 heavy-man\">Research Interests<\/p>\n<p class=\"pop-4\">My fundamental interests are in Plant Physiological Ecology and Biogeochemistry with special reference to trees and forest ecosystems. The theme that binds my interests together is an interest in how the microenvironment of a plant influences its form and function. In particular, I have been interested in how plants acclimate and adapt to environmental heterogeneity with an emphasis on how those physiological and morphological characteristics affect survival and growth. Recent projects have included stable isotopes in tree ring cellulose as indicators of plant water use and climate change, the effects of wind and leaf movements (leaf flutter) on canopy light dynamics and its impact on photosynthesis, the effects of elevated CO2 and temperature extreme events (global change) on tree seedling physiology.<\/p>\n<p>I am the faculty supervisor of the 麻豆传媒女神 Stable Isotope Facility (SOUSIF), where we measure the stable isotopes of C, O, and N from organic matter samples. Stable isotopes have numerous applications in biology and ecology and we have run samples for many different labs and investigators. Recently we had major NSF funded projects looking at the stable isotope variation in coast redwood trees as a proxy for climate variation over the last millennium.<\/p>\n<p class=\"dig-4 heavy-man\">Publications<\/p>\n<p class=\"pop-4\">Roden, J.S. (2008) Cross-dating of tree ring d18O and d13C time series. Chemical Geology. 252: 72-79. doi:10.1016\/j.chemgeo.2008.01.007<\/p>\n<p>Roden, J.S. Johnstone J., Dawson TE (2009) Intra-annual variation in the stable oxygen and carbon isotope ratios of cellulose in tree rings of coast redwood (Sequoia sempervirens). The Holocene. 19:189-197.<\/p>\n<p>Roden, JS, Canny, MJ, Huang CX and Ball, MC (2009) Frost tolerance and ice formation in Pinus radiata needles: ice management by the endodermis and transfusion tissues. Functional Plant Biology. 36:180\u2013189.<\/p>\n<p>Roden, J.S. Johnstone J.A. and T.E. Dawson (2011) Regional and watershed coherence in the stable oxygen and carbon isotope ratio time-series in tree rings of coast redwood (Sequoia sempervirens). Tree Ring Research. 67:71-86.<\/p>\n<p>Roden, JS and GD Farquhar (2012) A controlled test of the dual isotope approach for interpretation of stable carbon and oxygen isotope ratio variation in tree-rings. Tree Physiology. 32:490-503. doi:10.1093\/treephys\/tbs019.<\/p>\n<p>Roden, JS and R. Siegwolf (2012) Is the dual isotope conceptual model fully operational? Tree Physiology 32:1179-1182.<\/p>\n<p>Johnstone JA, J Roden, and TE Dawson (2013) Oxygen and carbon stable isotopes in coast redwood tree rings respond to spring and summer climate signals. Journal of Geophysical Research: Biogeosciences 118:1438-1450.<\/p>\n<p>Voelker SL, JR Brooks, FC Meinzer, J Roden, A Pazdur, S Pawelczyk, P Hartsough,K Snyder, L Plavcova and J \u0160antr\u016f\u010dek (2014) Reconstructing relative humidity from plant \u03b418O and \u03b4D as deuterium deviations from the global meteoric water line. Ecological Applications 24:960-975.<\/p>\n<p>Babst, F, Alexander R, Szejner P, Bouriaud O, Klesse S, Roden J, Ciais P, Poulter B, Frank D, Moore D and Trouet V. (2014) A tree-ring perspective on the terrestrial carbon cycle. Oecologia 176: 307-322. doi:10.1007\/s00442-014-3031-6<\/p>\n<p>Tipple BJ, Berke MA, Hambach, B, Roden JS and JR Ehleringer (2015) Predicting leaf wax n-alkane 2H\/1H ratios: controlled water source and humidity experiments with hydroponically grown trees confirm predictions of Craig\u2013Gordon model. Plant Cell and Environment 38:1035-1047. Doi:10.1111\/pce.12457.<\/p>\n<p>Roden JS, Kahmen A, Buchmann N and R Siegwolf (2015) The enigma of effective pathlength for 18O enrichment in leaf water of conifers. Plant Cell and Environment 38:2551-2565. doi:10.1111\/pce.12568.<\/p>\n<p>Churakova (Sidorova) O, Shashkin A, Launois T, Siegwolf R, Saurer M, Vaganov E, Bryukhanova M, Spahni R, Peylin P, Masson-Delmotte V and J Roden. Application of eco-physiological models to tree-ring parameters. 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